W Bateson and R C Punnett observed complementary gene interaction for flower colour In .)ݏݑݐܽݎ݀ ݏݑݎݕℎݐܽܮ) pea sweet in complementary interaction, two separate pairs of genes interact to produce the phenotype in such a way that neither of the dominant genes is expressive unless the other one is present. In F2 generation, complementary genes produce a ratio of 9 : 7.