Bridges gave genic balance theory of sex determination, which is related to ݏݎܦℎ݈݅ܽ ݈݉݁ܽ݊݃ܽݐݏ݁ݎ.According to him X-chromosome are carriers of genes for femaleness and autosomes are carriers of genes for maleness, so it is the ratio of X-chromosomes and autosomal sets which determine sex.