Hydropterids are only plant among the heterosporous. Pteridophytes that are leptosporangiate. Leptosporangiate in which the sporangium origin from epidermal cell Heterosporous pteridophytes were the first land flora of earth. The difference in size between microspore and megaspore is 1:2000 female gametophyte of Selaginella mostly have single archegonium