According to the active $K^{+}$ theory of Levitt, there is influx of $K^{+}$ in the guard cell from epidermal cells, which are balanced $by\ Cl^{-}$. At present, the role of $K^{+}$ in stomatal opening in universally accepted. In CAM plants, stomata remains closed during day time $e.g.,$ Opuntia, pineapple etc. Casparian strips present in the walls of endodermal cells control the amount of water and solute, which is absorbed, by the plants, CAM plants have capacity to store $CO_{2}$ during night and the same is used in photosynthesis during day time when stomata are closed